sentence: The involvement of sphingolipids in PCD ( Alden et al ., 2011 ; Liang et al ., 2003 ; Nagano et al ., 2009 ; Shi et al ., 2007 ) and in the response to pathogens ( Huby et al ., 2020 ; Wang et al ., 2008 ) by angiosperms may extend to bryophytes as well.Another , more subtle difference between sbh and WT colonies was observed : sbh protonemata often appeared drier than WT , which typically retain a large amount of moisture on the plant surface .