sentence: However , the extraction of plant material from herbarium specimens suitable for DNA analysis has been investigated with varying degrees of success depending on the technique used , age of material , preservation methods , and the conditions in which specimens have been stored ( e.g ., Rogers and Bendich , 1985 ; Liston et al ., 1990 ; Lister et al ., 2008 ; Andreasen et al ., 2009 ; S ä rkinen et al ., 2012 ; Kuzmina et al ., 2017 ) .