sentence: Apart from the the qFe5.1 , qFe6 , qAs4 , and qAs10 were overlapped with the Zinc-iron transporter gene OsZIP7a , OsZIP12 ( Yang et al ., 2009 ; Liu et al ., 2019 ) , and the arsenate reductase gene OsHAC1.2 and OsACR2.1 ( Duan et al ., 2007 ; Shi et al ., 2016 ) , respectively ( Table S2 ) , the other 29 QTLs found in O. longistaminata are different from the others .