sentence: Based on iWGS-based GWAS , we identified novel genomic regions of interest , revealing novel candidate genes ( ARHGAP35 , NPAS1 , TMEM160 , ZC3H4 , SAE1 , ZMIZ1 , PPIF , LDB2 , ABI3 , SERPINB6 , and SERPINB9 for LP ; NIM1K , ZNF131 , GABRG1 , GABRA2 , DCHS1 , and SPIDR for MILK ; NR6A1 , EXT2 , POLD1 , GOT1 , and ETV6 for FAT ; DPP6 , LRRC26 , and the KCN gene family for FAT % ; CDC14A , RTCA , HSTN , and ODAM for PROT ; and HERC3 , HERC5 , LALBA , and NEURL1 for PROT % ) , and confirmed previously reported associations .